What is the +2 and the +4? I thought those kind of runes did not stack?
Also if I already have a beastmastery rune on my armor piece and I then put a Marksmanship rune on it, does it over ride the Beastmastery rune?
The + 4 comes from a markmenship headpiece (+1) and a superior marksmenship (+3) :P
While the (+2) comes from a major rune of expertise.
Usually runing like this is pretty dangerous, and is often only done to reach breakpoints or for specific farming builds. The extra health is far more valuable than the small amount of damage you often get in return. Although this can be little more relaxed and you can often find rangers runing dual majors because of their better survivability (and because 14 expertise is awesome)
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Also if I already have a beastmastery rune on my armor piece and I then put a Marksmanship rune on it, does it over ride the Beastmastery rune?
No you can only have one rune per peice of armour :P
The question has been answered but to add a little...
Always put the highest attribute rune on a headpiece of the same attribute. That attribute should be the one you are going for the highest trained level in